[Illustration: Fig. 39. _a_, Paulia perforata, gonidia, paraphysis, and asci; _b_, Calicium tympanellum, perithecium and sporidia; _c_, Graphis Leprevostei, with excipulum, asci, and sporidium.]
The thallus or frond of many of these lichens is irregularly covered with thick convex scales, in each of which are concealed white or grey bodies, which become flask-shaped, with pores in their extremities. When mature, they exactly resemble the bottle-shaped perithecia in the genus Sphæria among fungi. They are lined with branching filaments or other supports, ending in minute ovoid particles, which escape in myriads from the flask-shaped vessels long before the spores appear. They are motile, though not provided with cilia, and are supposed to be fertilizing particles like pollen, since they do not germinate as spores do. In this vast family of open-shielded lichens, which comprises every form and habit of the plant, the preceding type, which is the highest, undergoes many modifications; but the essential character remains the same, and the group is so natural, that the species run into one another so as to leave no very striking distinction.
Commencing with the GYMNOCARPEI, the Parmeliacei first claim notice as containing the highest types which lichens are capable of assuming, and as abounding in species. The disc, which is the hymenium, is orbicular or kidney-shaped, and surrounded by the frond, without any proper excipulum or cortical covering. A large portion of our most widely diffused lichens, whether growing on the ground or attached to rocks and trees, belong to this tribe; they form patches two or three feet in diameter, which are often of marvellous antiquity, and they grow so slowly that even small patches are of great age. This tribe consists of three distinct groups: in two of these the plants are horizontal and sometimes foliaceous; in the third they are vertical, often branched, and occasionally pendulous.
The latter group of this large division of lichens comprises the genera allied to Usnea, which are erect, centripetal lichens, that is to say, their body or thallus is an upright and generally cylindrical stem, in the centre of which the layer of marrow, the origin of fructification, is condensed, and the open discs, or hymenia, are in connection with it, whether they be situated at the ends of the branches, or on the surface of the upright stems. The Usneæ have three forms, the pendulous, the inflated and erect, and the branched or shrubby. The Evernia jubata is an example of the first, and remarkable for its long, pendulous, cylindrical branches hanging down like bunches of bluish grey hair. It is often attached to the branches of aged larch trees, and is exceedingly picturesque. With the exception of Evernia vulpina and flavicans, which are brightly coloured, all the rest of the genus have dull tints. The genus Ramalina has the dull tints of the Evernia, and some species approach very nearly in form to that genus. The Ramalina polymorphum and R. scopulorum abound in dyes, while the R. calicaris is rich in gluten.