Pursuing this train of thought, I was led to think of the number of genera and species into which the plant world is divided,--a remarkable proof, not only of the power and wisdom, but of the goodness of the Creator, of His desire to furnish man with inexhaustible sources of pleasure and entertainment; and finally, to put to myself the question, How many vegetable species exist over the whole surface of the globe? If this corner of a leafy English lane is so rich in variety, what must be the case with "the wide, wide world?"
I was now brought to see that a question so difficult could, like so many others, be usefully approached only by its _inferior limit_; in other words, that in the actual condition of botanical science, we can but affirm the number which certainly _exceeds_ the sum of the vegetable species scattered over the surface of our earth. To determine this total with mathematical accuracy, we should need to have explored the terrestrial crust, liquid and solid, land and water, from the bed of ocean to the line of perpetual snow, and from the equator to the poles. And as yet we are very far from having obtained so complete a possession of the planet which has been assigned as a dwelling-place to our poor humanity,--alas, more presumptuous than powerful!
The number of plants mentioned by Theophrastus, Dioscorides, and Pliny, whom we take to be the representatives of ancient botany, does not exceed five hundred species. How very few, compared with the presumable total! The Middle Ages added scarcely anything to the botanical researches of antiquity. It is only since the discovery of America that we have seen the domain of Flora extending itself in unexpected proportions. But we must come down to the epoch of Linnæus (the middle of the eighteenth century) before we can obtain an accurate list of species, scientifically classified. Murray's edition of the "_Specilegium_" of Linnæus contains two thousand and forty-two species, including the Cryptogams. Wildmore, in another edition of the same great work, raised the total to twenty thousand. And this was the point at which our botanists had arrived when the nineteenth century opened.
But it was not long before they perceived that all these estimates, large as they seemed, fell immeasurably short of the reality. In attempting to distribute the different species among the then known regions of the globe, Alexander von Humboldt arrived at a total of forty-four thousand species, Phanerogams and Cryptogams included. De Candolle extended the estimate to upwards of fifty-six thousand.