THE PITH occupies the centre of the stem, and remains throughout the period of growth of some trees, as of the elder; or is abstracted after a few years, as in the oak, and almost all large trees. In the latter class of trees, there are some remains of the pith for many years after the process of absorption has commenced, but at length no vestige can be detected, and its position is known only by the central spot around which the wood is placed in circles. In the old age of the tree the pith frequently assumes a colour which it has obtained from the juices which have been deposited. The connections of the pith are extremely important. Firstly, it is in direct connection with every branch, and is the structure which first conveys fluids to, and receives fluids from every new leaf. It thence becomes the main organ of nutriment, and, at the same time, the chief depository of the secretions. Secondly, it is in equally direct and unbroken connection with the bark, through the medium of the medullary rays; and so becomes the centre of all the movements of sap which proceed in the horizontal system.
The mode in which the ultimate disappearance of the pith occurs has been a matter of speculation. That the circulation in the heart-wood ceases after a certain number of years, and that the connection between it and the bark becomes broken, is proved by the fact that numbers of trees may be found in tolerably vigorous growth within the bark, whereas at the heart they are decayed and rotten. It appears clear that it is not converted into wood, and there are facts against the opinion that it is gradually compressed by the wood; but since it is known that in the growth of the tree much compression of the previously formed wood must occur, and since this compression is a likely theory by which to account for the disappearance of the less resisting pith, it is now generally considered to be one of the causes of this occurrence. As a general rule, the pith, so long as it exists, is not mingled with other than cellular structures; but, in certain instances, wooden fibre has been found with it, and, in others, spiral vessels have been detected.
MEDULLARY SHEATH.--Immediately surrounding the pith of all exogenous plants, there is a layer of longitudinal tissue, which has received the name of medullary sheath. This sheath has no special walls, but is bounded by the pith on the inner, and the wood on the outer side. It is in this situation that ducts of various kinds and spiral vessels may be found, and in all cases it conveys the longitudinal structure from the root, direct to each leaf. The integrity of this structure is therefore highly necessary to the life of the tree.