Allosaurus () is an extinct genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period (Kimmeridgian to late Tithonian ages). The name "Allosaurus" means "different lizard", alluding to its unique (at the time of its discovery) concave vertebrae. It is derived from the Greek words () ("different", "strange", or "other") and () ("lizard" or "reptile"). The first fossil remains that could definitively be ascribed to this genus were described in 1877 by famed paleontologist Othniel Charles Marsh. As one of the first well-known theropod dinosaurs, it has long attracted attention outside of paleontological circles.
Allosaurus was a large bipedal predator for its time. Its skull was light, robust, and equipped with dozens of sharp, serrated teeth. It averaged in length for A. fragilis, with the largest specimens estimated as being long. Relative to the large and powerful legs, its three-fingered hands were small and the body was balanced by a long, muscular tail. It is classified as an allosaurid, a type of carnosaurian theropod dinosaur.
The genus has a very complicated taxonomy and includes at least three valid species, the best known of which is A. fragilis. The bulk of Allosaurus remains have come from North America's Morrison Formation, with material also known from the Lourinhã Formation in Portugal. It was known for over half of the 20th century as Antrodemus, but a study of the abundant remains from the Cleveland-Lloyd Dinosaur Quarry returned the name "Allosaurus" to prominence and established it as one of the best-known dinosaurs.
As the most abundant large predator of the Morrison Formation, Allosaurus was at the top of the food chain and probably preyed on contemporaneous large herbivorous dinosaurs, with the possibility of hunting other predators. Potential prey included ornithopods, stegosaurids, and sauropods. Some paleontologists interpret Allosaurus as having had cooperative social behavior and hunting in packs, while others believe individuals may have been aggressive toward each other and that congregations of this genus are the result of lone individuals feeding on the same carcasses.
Discovery and history
Early discoveries and research
The discovery and early study of Allosaurus is complicated by the multiplicity of names coined during the Bone Wars of the late 19th century. The first described fossil in this history was a bone obtained secondhand by Ferdinand Vandeveer Hayden in 1869. It came from Middle Park, near Granby, Colorado, probably from Morrison Formation rocks. The locals had identified such bones as "petrified horse hoofs". Hayden sent his specimen to Joseph Leidy, who identified it as half of a tail vertebra and tentatively assigned it to the European dinosaur genus Poekilopleuron as Poicilopleuron valens. He later decided it deserved its own genus, Antrodemus.
Allosaurus itself is based on YPM 1930, a small collection of fragmentary bones including parts of three vertebrae, a rib fragment, a tooth, a toe bone, and (most useful for later discussions) the shaft of the right humerus (upper arm). Othniel Charles Marsh gave these remains the formal name Allosaurus fragilis in 1877. Allosaurus comes from the Greek words /, meaning "strange" or "different", and /, meaning "lizard" or "reptile". It was named 'different lizard' because its vertebrae were different from those of other dinosaurs known at the time of its discovery. The species epithet fragilis is Latin for "fragile", referring to lightening features in the vertebrae. The bones were collected from the Morrison Formation of Garden Park, north of Cañon City. O. C. Marsh and Edward Drinker Cope, who were in scientific competition with each other, went on to coin several other genera based on similarly sparse material that would later figure in the taxonomy of Allosaurus. These include Marsh's Creosaurus and Labrosaurus, as well as Cope's Epanterias.