Granted, however, that a felt need of some kind, indefinite at first but none the less real, is present in many cases as a spur to instinctive behaviour; is it in all cases a necessary factor? May we say that this distinguishes instinctive from merely reflex action? The question is, from the nature of the case, exceedingly difficult to answer. But without going so far as to say that reflex action may be unerringly distinguished from instinctive behaviour by the absence of any such internal factors, we may perhaps, at any rate, go so far as to give provisional acceptance to the view that in instinct these wants and felt needs enter into the conscious situation in a manner and to a degree that are so far distinctive--which seems to be the position adopted by Professor Wundt.

There is, however, a further relation between the external stimulus and these internal factors which is presumably of no little importance. The stimulus intensifies the want, or may in some cases call it into existence. Just as a whiff from the kitchen may lead us to realize that we need the satisfaction that will erelong be presented at table, so may the sight of his mate in the spring evoke in the breast of the yearling sparrow a need, having its source in morphological and physiological changes, that spurs him on to the courtship that shall lead to its due satisfaction. Popular attention has, indeed, been so naturally drawn to the internal needs or wants with which we are now dealing, as to give them an almost exclusive monopoly of the term “instinct,” which thus often comes to be regarded as a connecting link between the stimulus and the act. The sight of a mouse, for example, is said to call forth the instinct of the cat, which is satisfied by her pouncing upon it. And so it comes about that, while the biologist fixes upon the instinctive act as the essential feature, the psychologist is apt to regard the impulse[41] which prompts to action as the more central and characteristic element. We are here endeavouring to combine both these points of view.

Animal Behaviour — Part 65

To come to closer quarters with the relationship which holds good between the external and internal elements, it appears that, when the stimulus evokes or intensifies the want or need, this is probably effected by efferent waves which call the organs or parts into tonic action, of which the animal becomes conscious through the afferent messages which come in from them to the sensory centres; in much the same way as the whiff from the kitchen takes effect on the salivary and other glands, and throws the organs of digestion into a felt preparedness for the fulfilment of their functions. But it may have other and more indirect consequences. When the moor-hen dived to escape from the obtrusive puppy, his heart-beat was probably affected; he had, perhaps, an uncomfortable sinking in his gizzard; his breathing was short and laboured; and he experienced creepy sensations in the skin and around the feather-roots. Such we may suppose were the accompaniments or sources of the emotional state of fear or alarm. And they presumably entered with no little vividness into the conscious situation at the moment of instinctive action. In all those cases in which the behaviour is associated with such an emotional state as anger or fear, the external stimulus seems to produce widely-spread effects on the glands, respiratory organs, heart and blood-vessels, skin and other parts, as well as the more direct response in productive action. And all this must enter into the conscious situation, contributing largely, as we shall hereafter see, to the emotions in their instinctive origin.

Animal Behaviour — Part 65