[Illustration: Fig. 6. Three Stages in the Development of an Exogenous Stem. P, pith; PB, primary bast; SB, secondary bast; C, cambium; PR, pith ray; PW, primary wood; SW, secondary wood; PS, procambium strands. _After Boulger._]
In the monocotyledons (endogens) these procambium strands change completely into wood and bast, and so losing all their protoplasmic cambium, become incapable of further growth. This is why palms can grow only lengthwise, or else by forming new fibers more densely in the central mass. But in the conifers and broad-leaved trees, the inner part of each strand becomes wood and the outer part bast (bark). Between these bundles, connecting the pith in the center with the cortex on the outside of the ring of bundles, are parts of the original pith tissue of the stem. They are the primary pith or medullary rays (Latin, _medulla_, pith). The number of medullary rays depends upon the number of the bundles; and their form, on the width of the bundles, so that they are often large and conspicuous, as in oak, or small and indeed invisible, as in some of the conifers. But they are present in all exogenous woods, and can readily be seen with the microscope. Stretching across these pith rays from the cambium layer in one procambium strand to that in the others, the cambium formation extends, making a complete cylindrical sheath from the bud downward over the whole stem. This is the cambium sheath and is the living, growing part of the stem from which is formed the wood on the inside and the rind (bark) on the outside.
In the first year the wood and the bast are formed directly by the growth and change of the inner and outer cells respectively of the procambium strand, and all such material is called "primary;" but in subsequent years all wood, pith rays, and bast, originate in the cambium, and these growths are called "secondary."
[Illustration: Fig. 7. Sap-wood and Heart-wood, Lignum Vitae.]
(3) The _wood_ of most exogens is made up of two parts, a lighter part called the sap-wood or splint-wood or alburnum, and a darker part called the heart-wood or duramen, Fig. 7. Sap-wood is really immature heartwood. The difference in color between them is very marked in some woods, as in lignum vitae and black walnut, and very slight in others, as spruce and bass. Indeed, some species never form a distinct heart-wood, birch (_Betula alba_) being an example.