If the relation of these cells to each other has been made sufficiently clear, it will be understood that, in accordance with Schewiakoff’s scheme of the structure, sections that cut the retinal cells transversely give very different appearances at different levels. A section through the very tops of the retinal cells, that is, the last section of the retina before striking the vitreous body, would show large polygonal areas of heavy pigment (the ends of the pigment cells), in between which would lie the much smaller, less pigmented, highly refracting ends of the visual cells (’89, Taf. II, Fig. 19). A section lower down in the retina, that is, more toward the centre of the club, would strike the low-lying enlarged central portion of the visual cells with their contained nuclei, and the smaller, proximal ends of the pigment cells. It would, therefore, give the reverse appearance from the preceding section, namely, that of large unpigmented (or but slightly pigmented) areas (the swollen bodies and nuclei of the spindle-shaped cells), and in between them smaller pigmented areas, the ends of the proximally tapering pigment cells (’89, Taf. II, Fig. 20). A section on the other side of the one first described, that is, one of the first through the vitreous body, would show pigment areas of the same size as the large ends of the pigment cells (the cone-shaped streaks of pigment in the vitreous body which according to Schewiakoff are associated with the pigment cell), and in between them the cross-sections of the rod-like processes from the visual cells, lying in canals in the clear homogeneous ground-substance of the vitreous body (’89, Taf. II, Fig. 18).

Let me give a resumé of Schewiakoff’s conception of the structure of the retina.

The Cubomedusæ — Part 46

a. There is an alternation of pigment and visual cells, the nuclei of the spindle-shaped visual cells lying at a lower level than those of the cone-shaped pigment cells.

The Cubomedusæ — Part 46

b. From the visual cells extend rod-like processes into the vitreous body, lying in canals in the latter.

The Cubomedusæ — Part 46

c. In the vitreous body a cone-shaped streak of pigment overlies each pigment cell of the retina, which is not a part of that cell.

d. Apart from these pigment streaks and the rod-like processes of the visual cells the vitreous body is structureless, probably a secretion of the pigment cells.