Even such an important and, among wild species, unusually constant organ as the bill has varied among pigeons to an astonishing degree. Carrier-pigeons (Fig. 1, No. 6) have an enormously long and strong bill, which is moreover covered with a thick red growth of the cere, while in the turbits and owls (Fig. 1, Nos. 8 and 10) the bill is shorter than any we find among wild birds. In many breeds even the _form_ of the bill deviates far from the normal, as in the bagadottes (No. 5) with crooked bill.

Like the bill, the legs vary in regard to their length. The pouters (No. 1) stand on their long legs as on stilts, while the legs of the 'Nürnberger swallow' are strikingly small. Remarkable, too, and very different from the wild species, is the thick growth of feathers on the feet and toes of the pouters and trumpeters (Fig. 1, No. 1), as well as of some other breeds, which suggests the arrangement of feathers on a wing.

The Evolution Theory, Vol. 1 of 2 — Part 40

Furthermore, the number and size of wing and tail-feathers in the different breeds often deviate considerably from the normal. The fantail (No. 7) in its most perfect form possesses forty tail-feathers, instead of the twelve usual in the wild rock-pigeon, and they are carried upright like a fan, while the head and neck of the bird are bent sharply backwards. In the hen-like pigeons the tail-feathers are few and short, so that they show an upright tail like that of a hen. I have already referred to the extraordinary carunculated skin-growth on the bill of many breeds; such folds also often surround the eye, and, as in the Indian barb (No. 3), are developed into well-formed thick circular ridges, while in the English carrier (No. 6) they lie about the bill as a formless mass of flesh.

The Evolution Theory, Vol. 1 of 2 — Part 40

Even the skull has undergone many variations, as can be observed even in the living bird in many of the breeds with short forehead. Differences are to be found, too, in the number and breadth of the ribs, the length of the breast-bone, the number and size of the tail-vertebræ in different breeds. Of the internal organs, the crop in many breeds, but particularly in the pouters (No. 1), has attained an enormous size, and with this size is usually associated the habit of blowing it out with air, and assuming the characteristically upright position.

The Evolution Theory, Vol. 1 of 2 — Part 40