For this reason, such an enormous increase as that of the caterpillars cannot last long; it carries its corrective in itself. The appearance of the caterpillars in such enormous numbers in itself increases the host of their enemies; singing-birds, ichneumon-flies, beetle-grubs, and predaceous beetles find abundant and available food, and therefore reproduce and multiply so rapidly, that, with the help of the caterpillar's plant-enemies, especially the insect-destroying fungi, they soon reduce the caterpillars to their normal number, or even below it. But then the reverse process begins; the enemies of the caterpillars diminish because their food has become scarce, and their normal number is lowered, while that of the caterpillars gradually rises again.

When the number of foxes in a hunting district increases, the number of the hares that they prey upon diminishes, and, on the other hand, the decimating of the foxes by Man brings about an increase in the number of hares in the district. Under natural conditions, that is, without the intervention of Man, there would be a constant balancing of the numbers of hares and foxes, for every noteworthy increase of the hares would be followed by a similar increase of foxes, and this, in its turn, would diminish the number of hares, so that they would no longer suffice for the support of so many foxes, and these would decrease in number again, until the number of hares had again increased because of the lessened persecution and elimination. In nature the case is not quite so simple, because the fox does not live on hares alone, and the hare is not preyed upon only by the fox; but the illustration may serve to elucidate the point that a moving equilibrium is maintained between the species of a district, between persecutors and persecuted, in such a way that the number of individuals in the two species is always varying a little up and down, and that each influences the other so that a regulative process results. Throughout periods of considerable length the average remains the same; that is to say, a _normal number_ is established. This normal strength of population is the mean above and below which the number of individuals is constantly varying. It is, of course, seldom that the mutual influences and regulations are so simple as in the example given; usually several or even many species interact upon each other, and not beasts of prey and their victims alone, but the most diverse species of animals and plants, which do not stand in any obvious relation to one another at all. Moreover, the physical, and especially the climatic conditions, also cause the normal number of the species to rise and fall.

The Evolution Theory, Vol. 1 of 2 — Part 51