The king penguin (Aptenodytes patagonicus) is the second largest species of penguin, smaller, but somewhat similar in appearance to the emperor penguin. There are two subspecies: A. p. patagonicus and A. p. halli; patagonicus is found in the South Atlantic and halli in the South Indian Ocean (at the Kerguelen Islands, Crozet Island, Prince Edward Islands, and Heard Island and McDonald Islands) and at Macquarie Island.
King penguins mainly eat lanternfish, squid, and krill. On foraging trips, king penguins repeatedly dive to over , and have been recorded at depths greater than . Predators of the king penguin include giant petrels, skuas, the snowy sheathbill, the leopard seal, and the orca.
The king penguin breeds on the Subantarctic islands at the northern reaches of Antarctica, South Georgia, southern Argentina, and other temperate islands of the region. It also lives on Macquarie Island in the Southern Ocean.
This bird was exploited commercially in the past for its blubber, oil, meat, and feathers. Today, it is fully protected.
Taxonomy
In 1778 the English illustrator John Frederick Miller included a hand-coloured engraving of the king penguin in his Icones animalium et plantarum. He coined the binomial name Aptenodytes patagonica and specified the type locality as the Mari antarctico, the Antarctic Ocean. The locality was restricted to South Georgia by Gregory Mathews in 1911. The species is monotypic: no subspecies are recognised.
Appearance
The king penguin stands at tall and weighs from . Although female and male king penguins are monomorphic they can be separated by their calls. Males are also slightly larger than females. The mean body mass of adults from Marion Island was for 70 males and for 71 females. Another study from Marion Island found that the mean mass of 33 adults feeding chicks was . The king penguin is approximately 25% shorter and weighs around a third less than the emperor penguin.
At first glance, the king penguin appears very similar to the larger, closely related emperor penguin, with a broad cheek patch contrasting with surrounding dark feathers and yellow-orange plumage at the top of the chest. However, the cheek patch of the adult king penguin is a solid bright orange whereas that of the emperor penguin is yellow and white, and the upper chest tends to be more orange and less yellowish in the king species. Both have colourful markings along the side of their lower mandible, but these tend towards pink in emperor penguin and orange in king penguin.
Emperor and king penguins typically do not inhabit the same areas in the wild, with the possible exception of vagrants at sea, but the two can be distinguished from one another by the king's longer, straighter bill, larger flippers, and noticeably sleeker body. The juvenile king penguin with its long bill and heavy dark brown down is completely different in appearance from the mostly grey emperor chick with its black and white mask. Once moulted of its brown juvenile plumage, the king chick resembles the adult, but is somewhat less colourful.
King penguins often breed on the same large, circumpolar islands as at least half of all living penguins, but it is easily distinguished from other species by its much larger size and taller frame, colorful markings, and grizzled sooty-greyish rather than blackish back.
Distribution and habitat
King penguins breed on subantarctic islands between 45 and 55°S, at the northern reaches of Antarctica, as well as Tierra del Fuego (Argentinian part), the Falkland Islands, and other temperate islands of the region. The total population is estimated to be 2.23 million pairs and is increasing. The largest breeding populations are on the Crozet Islands, with around 455,000 pairs, 228,000 pairs on the Prince Edward Islands, 240,000–280,000 on the Kerguelen Islands, and over 100,000 in the South Georgia archipelago. By the early 1920s, the King penguin population in South Georgia and the Falklands was nearly wiped out by whalers on these islands.
The Falklands and South Georgia had no trees to use for firewood, so the whalers burned millions of oily, blubber-rich penguins as fuel for the constant fires needed to boil whale blubber to extract the oil; the whalers also used penguin oil for lamps, heating, and cooking, in addition to eating the birds and their eggs. Macquarie Island currently has around 70,000 pairs. The non-breeding range is unknown due to many vagrant birds having been seen on the Antarctic peninsula as well as in South Africa, Australia, and New Zealand.