Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the food chain on plankton, often by filter feeding. They include particularly fishes of the order Clupeiformes (herrings, sardines, shad, hilsa, menhaden, anchovies, and sprats), but also other small fish, including halfbeaks, silversides, smelt such as capelin and goldband fusiliers.
Forage fish compensate for their small size by forming schools. Some swim in synchronised grids with their mouths open so they can efficiently filter plankton. These schools can become immense shoals which move along coastlines and migrate across open oceans. The shoals are concentrated energy resources for the great marine predators. The predators are keenly focused on the shoals, acutely aware of their numbers and whereabouts, and make migrations themselves that can span thousands of miles to connect, or stay connected, with them.
The ocean primary producers, mainly contained in plankton, produce food energy from the sun and are the raw fuel for the ocean food webs. Forage fish transfer this energy by eating the plankton and becoming food themselves for the top predators. In this way, forage fish occupy the central positions in ocean and lake food webs.
The fishing industry sometimes catch forage fish for commercial purposes, but primarily for use as feeder fish to farmed piscivorous animals. Some fisheries scientists are expressing concern that this will affect the populations of predator fish that depend on them.
In the oceans
Typical ocean forage fish are small, silvery schooling oily fish such as herring, anchovies and menhaden, and other small, schooling baitfish like capelin, smelts, sand lance, halfbeaks, pollock, butterfish and juvenile rockfish. Herrings are a preeminent forage fish, often marketed as sardines or pilchards.
The term “forage fish” is a term used in fisheries, and is applied also to forage species that are not true fish but play a significant role as prey for predators. Thus invertebrates such as squid and shrimp are also referred to as "forage fish". Even the tiny shrimp-like creatures called krill, small enough to be eaten by other forage fish, yet large enough to eat the same zooplankton as forage fish, are often classified as "forage fish".
Forage fish utilise the biomass of copepods, mysids and krill in the pelagic zone to become the dominant converters of the enormous ocean production of zooplankton. They are, in turn, central prey items for higher trophic levels. Forage fish may have achieved their dominance because of the way they live in huge, and often extremely fast cruising schools.
Though forage fish are abundant, there are relatively few species. There are more species of primary producers and apex predators in the ocean than there are forage fish.
Ocean food webs
Forage fish occupy central positions in the ocean food webs. The position that a fish occupies in a food web is called its trophic level (Greek trophē = food). The organisms it eats are at a lower trophic level, and the organisms that eat it are at a higher trophic level. Forage fish occupy middle levels in the food web, serving as a dominant prey to higher level fish, seabirds and mammals.
Ecological pyramids are graphical representations, along the lines of the diagram at the right, which show how biomass or productivity changes at each trophic level in an ecosystem. The first or bottom level is occupied by primary producers or autotrophs (Greek autos = self and trophe = food). These are the names given to organisms that do not feed on other organisms, but produce biomass from inorganic compounds, mostly by a process of photosynthesis.
In oceans, most primary production is performed by algae. This is a contrast to land, where most primary production is performed by vascular plants. Algae ranges from single floating cells to attached seaweeds, while vascular plants are represented in the ocean by groups such as the seagrasses. Larger producers, such as seagrasses and seaweeds, are mostly confined to the littoral zone and shallow waters, where they attach to the underlying substrate and are still within the photic zone. Most primary production in the ocean is performed by microscopic organisms, the phytoplankton.