On these grounds I designate the phenomenon as “polymorphism,” although it may not yet have reached, as such, its sharpest limits. This would be brought about by the elimination of the intermediate forms.[108]
Immediately before pupation, all the caterpillars, both green and brown, acquire a lilac coloration. The fifth stage lasts seven days, and the whole larval development twenty-three days, the period from the deposition of the eggs to the appearance of the moth being only thirty-one days.
I have treated of the polymorphism of _Stellatarum_ in detail, not only because it has hitherto remained unknown, and an analysis of such cases has been completely ignored,[109] but more particularly because, it appears to me, that important conclusions can be drawn therefrom. Moreover, such an extreme multiplicity of forms is interesting, since, so far as I know, polymorphism to this extent has not been observed in any insect.
The theoretical bearing of this polymorphism will be treated of subsequently. It is not in any way connected with a more advanced development of the markings, since _M. Stellatarum_ shows in this respect a very low state of development. This species displays only two stages:--(1), complete absence of all markings; and (2), a simple subdorsal, with dorsal and spiracular lines. We must therefore admit that the phyletic development of the markings has for a long time remained at a standstill, or, what expresses the same thing, that the marking which the adult larva now possesses is extremely old.
In order to complete my observations on _M. Stellatarum_, I now add some remarks on the pupa, the colour variations of which it appeared of importance to investigate, owing to the extraordinary variability of the caterpillar. The pupa varies but very slightly; the ochreous yellow ground-colour sometimes passes into reddish, and sometimes into greenish; the rather complicated blackish-brown marking of streaky lines is very constant, especially on the wing portions, being at most only more or less strongly pronounced. The minute colour variations of the pupa therefore have no connection with the colour of the caterpillar, both green and brown larvæ furnishing sometimes reddish-yellow and sometimes greenish-yellow pupæ.