Well over 100 players, coaches and owners throughout the National Football League defied U.S. President Donald Trump on Sunday by kneeling and linking arms in solidarity instead of standing when the national anthem was played at the beginning of their games.

The athletes said their protest is intended to draw attention to disparities in the treatment of racial minorities in the United States, including incidents of police brutality directed at African-Americans.

US Football Players Defy Trump, Protest During National Anthem

Trump has angrily denounced the protesters, and he repeated his disdain late Sunday, condemning the athletes for actions that he saw as "very disrespectful to our flag and to our country."

US Football Players Defy Trump, Protest During National Anthem

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US Football Players Defy Trump, Protest During National Anthem

To a reporter who asked if his own actions were inflaming racial tensions, Trump said: "No, this has nothing to do with race. I've never said anything about race."

Trump first raised the anthem issue at a political rally in Alabama on Friday night.

"Wouldn't you love to see one of these NFL owners, when somebody disrespects our flag, to say, 'Get that son of a bitch off the field right now. Out. He's fired. He's fired,'" Trump said. "Wouldn't you love it? Some owner's going to do that. He's going to say, 'That guy who disrespects our flag, he's fired.' And that owner ... - they're friends of mine, many of them - they'll be the most popular person for a week in this country."

Trump also suggested sports fans might boycott pro football games, which are among the most popular U.S. sporting events. The president backed off his earlier comments slightly as he headed back to the White House following a weekend at his golf course in New Jersey.

Asked specifically whether he thought protesting athletes should be fired, Trump said, "Well, I think the [team] owners should do something about it," but did not elaborate.

Roger Goodell, commissioner of the National Football League, which governs professional football in the U.S., said Trump's comments were divisive and disrespectful to athletes who were trying to make a heartfelt statement.

Asked to comment Sunday, Trump repeated his belief that those who knelt during the playing of the national anthem were "very disrespectful to our country ... very, very disrespectful to our flag."

Many athletes were bristling about the way in which Trump denounced the protests two days earlier during the Alabama rally.

Goodell's predecessor as NFL commissioner, Paul Tagliabue, said Trump's verbal assault was "disgraceful." Many athletes said they felt the president compounded his earlier unjustified insults by extending his disparaging comments to their families, and called for an apology.

Trump has not responded to those criticisms.

The football-field protests began last year as an individual action by an athlete who said he felt he had to make public his disapproval of the way in which racial minorities are treated in many parts of the U.S. The protests began to spread this year, and the salvo of tweets the president has fired off since Friday night have received wide publicity, and greatly swelled the numbers of players willing to speak out on the issue.

U.S. professional football games are mostly played on Sundays. The first game this week was in London, where the NFL is trying to build interest in American football. Many players, both black and white, knelt in protests when the music of "The Star Spangled Banner," the U.S. national anthem, was broadcast before play began between the Baltimore Ravens and Jacksonville Jaguars.

All of the players stood when the British anthem, "God Save the Queen," resounded through the stadium, as is customary before an athletic contest.

Not only individual football stars, but other team officials and owners have joined the fray, and they are not siding with the president.

Robert Kraft, owner of the defending Super Bowl champion New England Patriots, is a Trump supporter who donated $1 million to the president's inaugural committee earlier this year. He said he was "deeply disappointed by the tone of the comments" Trump made.

"I am proud to be associated with so many players who make such tremendous contributions in positively impacting our communities," said Kraft, who has visited with Trump at the White House. "Our players are intelligent, thoughtful and care deeply about our community, and I support their right to peacefully affect social change and raise awareness in a manner that they feel is most impactful."