A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest, is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud cover, usually at the canopy level, formally described in the International Cloud Atlas (2017) as silvagenitus. Cloud forests often exhibit an abundance of mosses covering the ground and vegetation, in which case they are also referred to as mossy forests. Mossy forests usually develop on the saddles of mountains, where moisture introduced by settling clouds is more effectively retained.
Cloud forests are among the most biodiversity rich ecosystems in the world with a large number of species directly or indirectly depending on them.
Other moss forests include black spruce/feathermoss climax forest, with a moderately dense canopy and a forest floor of feathermosses including Hylocomium splendens, Pleurozium schreberi and Ptilium crista-castrensis. These weft-form mosses grow in boreal moss forests.
Climate
The presence of cloud forests is dependent on local climate (which is affected by the distance to the sea), the exposition and the latitude (from 23°N to 25°S), and the elevation (which varies from 500 m to 4000 m above sea level). Typically, there is a relatively small band of elevation in which the atmospheric environment is suitable for cloud forest development. This is characterized by persistent fog at the vegetation level, resulting in the reduction of direct sunlight and thus of evapotranspiration. Within cloud forests, much of the moisture available to plants arrives in the form of fog drip, where fog condenses on tree leaves and then drips onto the ground below.
Annual rainfall can range from 500 to 10,000 mm/year and mean temperature between 8 and 20 °C (46.4 and 68 °F).
While cloud forest today is the most widely used term, in some regions, these ecosystems or special types of cloud forests are called mossy forest, elfin forest, montane thicket, and dwarf cloud forest.
The definition of cloud forest can be ambiguous, with many countries not using the term (preferring such terms as Afromontane forest and upper montane rain forest, montane laurel forest, or more localised terms such as the Bolivian yungas, and the laurisilva of the Atlantic Islands), and occasionally subtropical and even temperate forests in which similar meteorological conditions occur are considered to be cloud forests.
Characteristics
In comparison with lower-altitude tropical moist forests, cloud forests show a reduced tree stature combined with increased stem density and generally, a lower diversity of woody plants. Trees in these regions are generally shorter and more heavily stemmed than in lower-altitude forests in the same regions, often with gnarled trunks and branches, forming dense, compact crowns. Their leaves become smaller, thicker and harder with increasing altitude. The high moisture promotes the development of a high biomass and biodiversity of epiphyte, particularly bryophytes, lichens, ferns (including filmy ferns), bromeliads and orchids. The number of endemic plants can be very high.
An important feature of cloud forests is the tree crowns that intercept the wind-driven cloud moisture, part of which drips to the ground. This fog drip occurs when water droplets from the fog adhere to the needles or leaves of trees or other objects, coalesce into larger drops and then drop to the ground. It can be an important contribution to the hydrologic cycle.
Cloud forests are often peatlands, showcasing many classic peatland attributes. Due to the high water content of the soil, the reduced solar radiation and the low rates of decomposition and mineralization, the soil acidity is very high, with more humus and peat often forming the upper soil layer.
Stadtmüller (1987) distinguishes two general types of tropical montane cloud forests: Areas with a high annual precipitation due to a frequent cloud cover in combination with heavy and sometimes persistent orographic rainfall; such forests have a perceptible canopy strata, a high number of epiphytes, and a thick peat layer which has a high storage capacity for water and controls the runoff; In drier areas with mainly seasonal rainfall, cloud stripping can amount to a large proportion of the moisture available to plants.