Snails have existed from the earliest Paleozoic era to the present time, and the outstanding fact of interest concerning them is that they furnish one of the finest illustrations of an important class of animals which has undergone practically no conspicuous change or evolution during all those millions of years of time.
[Illustration: Fig. 53.--Sketches of chambered cephalopods showing the main steps in the evolution of the shell forms and compartment partitions: a, b, the only kinds in Cambrian time; c, d, forms added in the Ordovician; e, added in the Devonian; f, added in the late Paleozoic; g, h, characteristic of the Mesozoic era; and i, a living form (pearly nautilus) cut through. (Drawn by the author.)]
We shall now turn our attention to the highest order of Mollusks--the cephalopods. These creatures, whose heads are armed with powerful tentacles and supplied with complex eyes, propel themselves by forcible ejection of water. One general type--the chambered cephalopod--has a shell divided into compartments (e.g., modern pearly nautilus) which are successively built up and abandoned by the animal as it grows larger. These chamber-shelled cephalopods constitute one of the most remarkable and instructive illustrations of evolutionary change within any important subgroup of invertebrate animals, ranging from early Paleozoic to the present. Both because of the abundance of fossil forms in rocks of all these periods of geological times, and because certain of the evolutionary changes are so clearly expressed in the well preserved shell portions, they are specially adapted for study. In the late Cambrian only straight and slightly curved forms with smooth, nearly straight chamber partitions existed. Notable advance took place during the next (Ordovician) period when there were straight, curved, open-coiled, and even close-coiled forms. All had simple partitions, and the straighter forms predominated. "The size attained by the Ordovician cephalopods was probably never surpassed by representatives of the class. Some of the (straight) shells were twelve to fifteen feet in length, and a foot in diameter. From this great size they ranged down to or below the size of a pipe stem." (Chamberlin and Salisbury.) They were more than likely the undisputed masters of the Ordovician seas. Silurian time marked no important change in their structures, but the coiled forms predominated for the first time. During the second half of the Paleozoic era all preceding types with simple partitions persisted, but in some forms the simple partitions gradually became angled and finally rather complexly curved. During the Mesozoic era the partition lines of the close-coiled forms evolved until a most remarkable degree of complexity was attained, comparable, indeed, to the sutures of the human skull plates. These remarkable forms called ammonites, of which more than 2,000 species are known, began with the Mesozoic, reached their climax, and passed out of existence toward the close of the same era. Certain strata of Jurassic age are literally filled with ammonites, some shells being several feet in diameter. Various eccentric changes took place in the ammonites shortly before their extinction. Some shells became uncoiled and even straight, thus outwardly at least showing reversion to the original early Paleozoic ancestors, but with retention of the complex partitions. Others assumed spiral shapes and still others became curved or coiled at each end. While these extraordinary evolutionary changes were going on among the chambers of cephalopods during Mesozoic time, some of the ancient close-coiled forms with very simple partitions managed to persist. In fact this simple type, almost exactly like its early Paleozoic ancestor, has been the only one out of this whole remarkable class of animals to persist to the present time, being now barely represented by the well-known pearly nautilus of the Indian Ocean.