If [this] law be true, it follows that the natural series of affinities will also represent the order in which the several species came into existence, each one having had for its immediate antetype a clearly allied species existing at the time of its origin.... If two or more species have been independently formed on the plan of a common antetype, then the series of affinities will be compound, and can only be represented by a forked or many-branched line.... Sometimes the series of affinities can be well represented for a space by a direct progression from species to species or from group to group, but it is generally found impossible so to continue. There constantly occur two or more modifications of an organ or modifications of two distinct organs, leading us on to two distinct series of species, which at length differ so much from each other as to form distinct genera or families. These are the parallel series or representative groups of naturalists, and they often occur in different countries, or are found fossil in different formations.... We thus see how difficult it is to determine in every case whether a given relation is an analogy or an affinity, for it is evident that as we go back along the parallel or divergent series, towards the common antetype, the analogy which existed between the two groups becomes an affinity.... Again, if we consider that we have only the fragments of this vast system, the stems and main branches being represented by extinct species of which we have no knowledge, while a vast mass of limbs and boughs and minute twigs and scattered leaves is what we have to place in order, and determine the true position each originally occupied with regard to the others, the whole difficulty of the true Natural System of classification becomes apparent to us.
We shall thus find ourselves obliged to reject all those systems of classification which arrange species or groups in circles, as well as those which fix a definite number for the division of each group.... We have ... never been able to find a case in which the circle has been closed by a direct affinity. In most cases a palpable analogy has been substituted, in others the affinity is very obscure or altogether doubtful....