I agree with Dr. Williamson[U] in believing that all or nearly all the forms referred to Crossochorda of Schimper are really animal impressions allied to Nereites, and due either to worms or, as Nathorst has shown to be possible, to small crustaceans. Many impressions of this kind occur in the Silurian beds of the Clinton series in Canada and New York, and are undoubtedly mere markings.
[U] "Tracks from Yoredale Rocks," "Manchester Literary and Philosophical Society," 1885.
[Illustration: Fig. 9.--_Astropolithon Hindii_, an organism of the Lower Cambrian of Nova Scotia, possibly vegetable.]
It is worthy of note that these markings strikingly resemble the so-called _Eophyton_, described by Torell from the Primordial of Sweden, and by Billings from that of Newfoundland; and which also occur abundantly in the Primordial of New Brunswick. After examining a series of these markings from Sweden shown to me by Mr. Carruthers in London, and also specimens from Newfoundland and a large number _in situ_ at St. John, I am convinced that they cannot be plants, but must be markings of the nature of _Rhabdichnites_. This conclusion is based on the absence of carbonaceous matter, the intimate union of the markings with the surface of the stone, their indefinite forms, their want of nodes or appendages, and their markings being always of such a nature as could be produced by scratches of a sharp instrument. Since, however, fishes are yet unknown in beds of this age, they may possibly be referred to the feet or spinous tails of swimming crustaceans. Salter has already suggested this origin for some scratches of somewhat different form found in the Primordial of Great Britain. He supposed them to have been the work of species of _Hymenocaris_. These marks may, however, indicate the existence of some free-swimming animals of the Primordial seas as yet unknown to us.
Three other suggestions merit consideration in this connection. One is that Algæ and also land-plants, drifting with tides or currents, often make the most remarkable and fantastic trails. A marking of this kind has been observed by Dr. G. M. Dawson to be produced by a drifted Laminaria, and in complexity it resembled the extraordinary _Ænigmichnus multiformis_ of Hitchcock from the Connecticut sandstones. Much more simple markings of this kind would suffice to give species of _Eophyton_. Another is furnished by a fact stated to the author by Prof. Morse, namely, that Lingulæ, when dislodged from their burrows, trail themselves over the bottom like worms, by means of their cirri. Colonies of these creatures, so abundant in the Primordial, may, when obliged to remove, have covered the surfaces of beds of mud with vermicular markings. The third is that the Rhabdichnite-markings resemble some of the grooves in Silurian rocks which have been referred to trails of Gasteropods, as, for instance, those from the Clinton group, described by Hall.