The larger wild animals are not now numerous enough in any one region to form extensive deposits by their remains; but they have, nevertheless, a certain geographical importance. If the myriads of large browsing and grazing quadrupeds which wander over the plains of Southern Africa--and the slaughter of which by thousands is the source of a ferocious pleasure and a brutal triumph to professedly civilized hunters--if the herds of the American bison, which are numbered by hundreds of thousands, do not produce visible changes in the forms of terrestrial surface, they have at least an immense influence on the growth and distribution of vegetable life, and, of course, indirectly upon all the physical conditions of soil and climate between which and vegetation a mutual interdependence exists.
The influence of wild quadrupeds upon vegetable life has been little studied, and not many facts bearing upon it have been recorded, but, so far as it is known, it appears to be conservative rather than pernicious.[61] Few if any of them depend for their subsistence on vegetable products obtainable only by the destruction of the plant, and they seem to confine their consumption almost exclusively to the annual harvest of leaf or twig, or at least of parts of the vegetable easily reproduced. If there are exceptions to this rule, they are in cases where the numbers of the animal are so proportioned to the abundance of the vegetable, that there is no danger of the extermination of the plant from the voracity of the quadruped, or of the extinction of the quadruped from the scarcity of the plant. In diet and natural wants the bison resembles the ox, the ibex and the chamois assimilate themselves to the goat and the sheep; but while the wild animal does not appear to be a destructive agency in the garden of nature, his domestic congeners are eminently so. This is partly from the change of habits resulting from domestication and association with man, partly from the fact that the number of reclaimed animals is not determined by the natural relation of demand and spontaneous supply which regulates the multiplication of wild creatures, but by the convenience of man, who is, in comparatively few things, amenable to the control of the merely physical arrangements of nature. When the domesticated animal escapes from human jurisdiction, as in the case of the ox, the horse, the goat, and perhaps the ass--which, so far as I know, are the only well-authenticated instances of the complete emancipation of household quadrupeds--he becomes again an unresisting subject of nature, and all his economy is governed by the same laws as that of his fellows which have never been enslaved by man; but, so long as he obeys a human lord, he is an auxiliary in the warfare his master is ever waging against all existences except those which he can tame to a willing servitude.