In _Sphenodon_, then, on the dorsal surface of the skull, are the large =supratemporal fossae= (fig. 52, 20). Their inner margins are separated from one another by the parietal walls of the cranium, while externally each is bounded by a bony arch, the =supratemporal arcade=, formed of the postfrontal, postorbital, and squamosal. Posteriorly the boundary is formed by a =post-temporal bar=, formed by the parietal and squamosal. Below the supratemporal arcade is another large vacuity, the =infratemporal= or =lateral temporal fossa= (fig. 52, 21). This is bounded above by the supratemporal arcade, and is separated from the orbit in front by the =postorbital bar=, formed by the union of outgrowths from the jugals and postorbitals. Behind it is bounded by a continuation of the post-temporal bar formed of the squamosal and quadratojugal, and below by an =infratemporal arcade=, which is chiefly composed of the quadratojugal and jugal.

Below the post-temporal bar is a third vacuity, the =post-temporal fossa= (fig. 52, D, 23), bounded above by the post-temporal bar and below by the exoccipital and opisthotic.

The Vertebrate Skeleton — Part 176

_Sphenodon_ and the Crocodilia are the only living reptiles with complete supratemporal and infratemporal arcades, but they are both present in the extinct Pterosauria and some Dinosauria.

The Vertebrate Skeleton — Part 176

Supratemporal fossae, bounded below by supratemporal arcades, occur in all reptiles except some Chelonia, the Ophidia, the Geckonidae among Lacertilia, and the Pariasauria and others among Theromorpha; they are specially large in _Nothosaurus_ among the Sauropterygia, _Dicynodon_ among the Theromorpha, and many Crocodilia and Pterosauria. In some Dinosaurs, such as _Ceratosaurus_, they are very small, while the infratemporal fossae are correspondingly large.

The Vertebrate Skeleton — Part 176

In _Elginia_[93] (Theromorpha) and some Chelonia, such as _Chelone_, there are no fossae on the surface of the skull, a complete false roof being developed; in other Chelonia, such as _Trionyx_, the true cranium is freely visible, the only part of the false roof developed being the infratemporal arcade.

In many reptiles large =pre-orbital vacuities= occur; they are specially large in the Pterosauria and in some of the Crocodilia and Dinosauria (fig. 35, 3). In some Pterosauria they are confluent with the orbits.