[39] It perhaps deserves notice that besides these five birds two of the eight were Barbs, which, as I have shown, must be classed in the same group with the long-beaked Carriers and Runts. Barbs may properly be called short-beaked Carriers. It would, therefore, appear as if, during the reduction of their beaks, their wings had retained a little of that excess of length which is characteristic of their nearest relations and progenitors.

[40] Temminck, ‘Hist. Nat. Gén. des Pigeons et des Gallinacés,’ tom. i., 1813, p. 170.

The Variation of Animals and Plants under Domestication — Part 166

[41] This term was used by John Hunter for such differences in structure between the males and females, as are not directly connected with the act of reproduction, as the tail of the peacock, the horns of deer, etc.

The Variation of Animals and Plants under Domestication — Part 166

ON THE ABORIGINAL PARENT-STOCK OF THE SEVERAL DOMESTIC RACES—HABITS OF LIFE—WILD RACES OF THE ROCK-PIGEON—Dovecot-PIGEONS—PROOFS OF THE DESCENT OF THE SEVERAL RACES FROM COLUMBA LIVIA—FERTILITY OF THE RACES WHEN CROSSED—REVERSION TO THE PLUMAGE OF THE WILD ROCK-PIGEON—CIRCUMSTANCES FAVOURABLE TO THE FORMATION OF THE RACES—ANTIQUITY AND HISTORY OF THE PRINCIPAL RACES—MANNER OF THEIR FORMATION—SELECTION—UNCONSCIOUS SELECTION—CARE TAKEN BY FANCIERS IN SELECTING THEIR BIRDS—SLIGHTLY DIFFERENT STRAINS GRADUALLY CHANGE INTO WELL-MARKED BREEDS—EXTINCTION OF INTERMEDIATE FORMS—CERTAIN BREEDS REMAIN PERMANENT, WHILST OTHERS CHANGE—SUMMARY.

The Variation of Animals and Plants under Domestication — Part 166

The differences described in the last chapter between the eleven chief domestic races and between individual birds of the same race, would be of little significance, if they had not all descended from a single wild stock. The question of their origin is therefore of fundamental importance, and must be discussed at considerable length. No one will think this superfluous who considers the great amount of difference between the races, who knows how ancient many of them are, and how truly they breed at the present day. Fanciers almost unanimously believe that the different races are descended from several wild stocks, whereas most naturalists believe that all are descended from the Columba livia or rock-pigeon.

Temminck[1] has well observed, and Mr. Gould has made the same remark to me, that the aboriginal parent must have been a species which roosted and built its nest on rocks; and I may add that it must have been a social bird. For all the domestic races are highly social, and none are known to build or habitually to roost on trees. The awkward manner in which some pigeons, kept by me in a summer-house near an old walnut-tree, occasionally alighted on the barer branches, was evident.[2] Nevertheless, Mr. R. Scot Skirving informs me that he often saw crowds of pigeons in Upper Egypt settling on low trees, but not on palms, in preference to alighting on the mud hovels of the natives. In India Mr. Blyth[3] has been assured that the wild _C. livia,_ var. _intermedia,_ sometimes roosts in trees. I may here give a curious instance of compulsion leading to changed habits: the banks of the Nile above lat. 28° 30′ are perpendicular for a long distance, so that when the river is full the pigeons cannot alight on the shore to drink, and Mr. Skirving repeatedly saw whole flocks settle on the water, and drink whilst they floated down the stream. These flocks seen from a distance resembled flocks of gulls on the surface of the sea.