The condition of the several parts of the skeleton completely proves that the Pterodactyle was a reptile. Its head, neck, shoulder, and back, resemble a bird; while there are, on the other hand, some striking resemblances with the reptile in the pelvis, tail, and articular parts of the limbs. Sometimes the characters of the two classes run side by side, as in the skull, the fore-limbs, and especially in the hind-limbs, where the shin of a Bird is connected with the foot of a Saurian. The parts in which it corresponds with birds show that Pterodactyles also were flying animals. That we should be entitled to conclude, from the hollow state of the bones, that they belonged to flying animals, is sufficiently proved by Blumenbach, Buckland, Mantell, Owen having mistaken them for bones of birds.

The most absolute proof that it was a flying animal is the pneumatic character of its bones. This condition was discerned by me in some Pterodactyle bones from the Lias of Franken (_Jahrb. für Mineral_, 1837, p. 316), and was afterwards established by Owen in the Pterodactyles from the Chalk of England. This structure was previously only known in birds. And the supposition readily follows that in the respiratory process there was some similarity between the Pterodactyle and the Birds. They have the proportions of upper-arm and fore-arm which characterize birds of great flight, the humerus short and the fore-arm long; hence it may be presumed that Pterodactyles could fly well. From the absence and presence of the bony sclerotic ring in the eye, it may be supposed that the Pterodactyles were active in the day-time, while Rhamphorhynchus was nocturnal.

The Ornithosauria: an elementary study of the bones of pterodactyles — Part 19

After this statement von Meyer gives a discursive summary, in which his views of the classification of reptiles in general and of Pterodactyles in particular are epitomized. And then goes on to combat the views of people who have departed from his classification and attempted to set up classifications of their own; and cites a number of authors who, labouring at the vertebrata, have endeavoured to find a resting-place in their systems for the Pterodactyle. But the chief thing we learn of von Meyer's own views is, that in 1830 he published a classification of extinct Saurians, dividing them into those with limbs like the larger and heavier land-mammals, those with fin-like limbs, and those with a flying-finger. Which divisions have been widely adopted, though authors have sometimes given them other names than those by which they were first made known.

The Ornithosauria: an elementary study of the bones of pterodactyles — Part 19