Perhaps one cannot more forcibly present the rational value of this position than by noting the following consequences of it. Mr. Gulick writes me that while studying the land-shells of the Sandwich Islands, and finding there a rich profusion of unique varieties, in cases where the intermediate varieties were rare he could himself have created a number of species by simply throwing these intermediate varieties into his fire. Now it follows from the dogma which we are considering, that, by so doing, not only would he have created new species, but at the same time he would have proved them due to natural selection, and endowed the diagnostic characters of each with a "necessarily" adaptive meaning, which previously it was not necessary that they should present. Before his destruction of these intermediate varieties, he need have felt himself under no obligation to assume that any given character at either end of the series was of utilitarian significance: but, after his destruction of the intermediate forms, he could no longer entertain any question upon the matter, under pain of being denounced as a Darwinian heretic.
Now the application is self-evident. It is a general fact, which admits of no denial, that the more our knowledge of any flora or fauna increases, the greater is the number of intermediate forms which are brought to light, either as still existing or as having once existed. Consequently, the more that such knowledge increases, the more does our catalogue of "species" diminish. As Kerner says, "bad species" are always multiplying at the expense of "good species"; or, as Oscar Schmidt (following Häckel) similarly remarks, if we could know as much about the latter as we do about the former, "all species, without any exception, would become what species-makers understand by 'bad species'[129]." Hence we see that, just as Mr. Gulick could have created good species by secretly destroying his intermediate varieties, so has Nature produced her "good species" for the delectation of systematists. And just as Mr. Gulick, by first hiding and afterwards revealing his intermediate forms, could have made the self-same characters in the first instance necessarily useful, but ever afterwards presumably useless, so has Nature caused the utility of diagnostic characters to vary with our knowledge of her intermediate forms. It belongs to the essence of our theory of descent, that in _all_ cases these intermediate forms must either be now existing or have once existed; and, therefore, that the work of species-makers consists in nothing more than marking out the _lacunae_ in our knowledge of them. Yet we are bound to believe that wherever these _lacunae_ in our knowledge occur, there occurs also the objective necessity of causation as utilitarian--a necessity, however, which vanishes so soon as our advancing information supplies the intermediate forms in question. It may indeed appear strange that the utility or non-utility of organic structures should thus depend on the accidents of human knowledge; but this is the Darwinian faith, and he who doubts the dogma is to be anathema.