The vast mass of material made known by these and many other distinguished writers has to be included in our classification, and that classification itself must be controlled by the story it reveals. Thus it is that a change, characteristic of modern systematic zoology, is affecting the subdivisions of the classes. It is not long since the main lines of division corresponded roughly to gaps in geological history: the orders were Palaeocrinoidea and Neocrinoidea, Palechinoidea and Euechinoidea, Palaeasteroidea and Euasteroidea, and so forth. Or divisions were based upon certain modifications of structure which, as we now see, affected assemblages of diverse affinity: thus both Blastoidea and Euechinoidea were divided into Regularia and Irregularia; the Holothuroidea into Pneumophora and Apneumona; and Crinoids were discussed under the heads "stalked" and "unstalked." The barriers between these groups may be regarded as horizontal planes cutting across the branches of the ascending tree of life at levels determined chiefly by our ignorance; as knowledge increases, and as the conception of a genealogical classification gains acceptance, they are being replaced by vertical partitions which separate branch from branch. The changes may be appreciated by comparing the systematic synopses at the end of this article with the classification adopted in 1877 in the 9th edition of the _Ency. Brit._ (vol. vii.), or in any zoological text-book contemporary therewith. In the present stage of our knowledge these minor divisions are the really important ones. For, whereas to one brilliant suggestion of far-reaching homology another can always be opposed, by the detailed comparison of individual growth-stages in carefully selected series of fossils, and by the minute application to these of the principle that individual history repeats race history, it actually is possible to unfold lines of descent that do not admit of doubt. The gradual linking up of these will manifest the true genealogy of each class, and reconstruct its ancestral forms by proof instead of conjecture. The problem of the interrelations of the classes will thus be reduced to its simplest terms, and even questions as to the nature of the primitive Echinoderm and its affinity to the ancestors of other phyla may become more than exercises for the ingenuity of youth. Work has been and is being done by the laborious methods here alluded to, and though the diversity of opinion as to the broader groupings of classification is still restricted only by the number of writers, we can point to an ever-increasing body of assured knowledge on which all are agreed. Unfortunately such allusion to these disconnected certainties as alone might be introduced here would be too brief for comprehension, and we are forced to select a few of the broader hypotheses for a treatment that may seem dogmatic and prejudiced.
Frontiers Report
Encyclopaedia Britannica, 11th Edition, Volume 8, Slice 10 — Part 4
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