The maxillulae and maxillae (or, as they are often termed, first and second maxillae) are nearly always flattened leaf-like appendages, having gnathobasic lobes or endites borne by the segments of the protopodite. The endopodite, when present, is unsegmented or composed of few segments and forms the "palp," and outwardly-directed lobes representing the exopodite and epipodites may also be present. These limbs undergo great modification in the different groups. The maxillulae are sometimes closely connected with the "paragnatha" or lobes of the lower lip, when these are present, and it has been suggested that the paragnatha are really the basal endites which have become partly separated from the rest of the appendage.

The limbs of the post-cephalic series show little differentiation among themselves in many Entomostraca. In the Phyllopoda they are for the most part all alike, though one or two of the anterior pairs may be specialized as sensory (_Apus_) or grasping (_Estheriidae_) organs. In the Cirripedia (Thyrostraca) the six pairs of biramous cirriform limbs differ only slightly from each other, and in many Copepoda this is also the case. In other Entomostraca considerable differentiation may take place, but the series is never divided into definite "tagmata" or groups of similarly modified appendages. It is highly characteristic of the Malacostraca, however, that the trunk-limbs are divided into two sharply defined tagmata corresponding to the thoracic and abdominal regions respectively, the limit between the two being marked by the position of the male genital openings. The thoracic limbs have the endopodites converted, as a rule, into more or less efficient walking-legs, and the exopodites are often lost, while the abdominal limbs more generally preserve the biramous form and are, in the more primitive types, natatory. These tagmata may again be subdivided into groups preserving a more or less marked individuality. For example, in the Amphipoda (fig. 4) the abdominal appendages are constantly divided into an anterior group of three natatory "swimmerets" and a posterior group of three limbs used chiefly in jumping or in burrowing. In nearly all Malacostraca the last pair of abdominal appendages (uropods) differ from the others, and in the more primitive groups they form, with the telson, a lamellar "tail-fan" (fig. 3, T), used in springing backwards through the water. In the thoracic series it is usual for one or more of the anterior pairs to be pressed into the service of the mouth, forming "foot-jaws" or maxillipeds. In the Decapoda three pairs are thus modified, and in the Tanaidacea, Isopoda and Amphipoda only one. In the Schizopoda and Cumacea the line of division is less sharp, and the varying number of so-called maxillipeds recognized by different authors gives rise to some confusion of terminology in systematic literature.

Encyclopaedia Britannica, 11th Edition, Volume 7, Slice 7 — Part 192